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Anatomy is the study of internal structure of organism. Study of plant anatomy includes histology- study of organization and structure of tissues. Anatomy helps in knowing the structural peculiarities of different group of plants and indicates the structural adaptation to diverse environments.
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A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). Flowers produce gametophytes, which in flowering plants consist of a few haploid cells which produce gametes. The "male" gametophyte, which produces non-motile sperm, is enclosed within pollen grains; the "female" gametophyte is contained within the ovule. When pollen from the anther of a flower is deposited on the stigma, this is called pollination. Some flowers may self-pollinate, producing seed using pollen from the same flower or a different flower of the same plant, but others have mechanisms to prevent self-pollination and rely on cross-pollination, when pollen is transferred from the anther of one flower to the stigma of another flower on a different individual of the same species.
A flower develops on a modified shoot or axis from a determinate apical meristem (determinate meaning the axis grows to a set size). It has compressed internodes, bearing structures that in classical plant morphology are interpreted as highly modified leaves.[33] Detailed developmental studies, however, have shown that stamens are often initiated more or less like modified stems (caulomes) that in some cases may even resemble branchlets.[34][15] Taking into account the whole diversity in the development of the androecium of flowering plants, we find a continuum between modified leaves (phyllomes), modified stems (caulomes), and modified branchlets (shoots).[35][36]
The transition to flowering is one of the major phase changes that a plant makes during its life cycle. The transition must take place at a time that is favorable for fertilization and the formation of seeds, hence ensuring maximal reproductive success. To meet these needs a plant is able to interpret important endogenous and environmental cues such as changes in levels of plant hormones and seasonable temperature and photoperiod changes.[37] Many perennial and most biennial plants require vernalization to flower. The molecular interpretation of these signals is through the transmission of a complex signal known as florigen, which involves a variety of genes, including Constans, Flowering Locus C, and Flowering Locus T. Florigen is produced in the leaves in reproductively favorable conditions and acts in buds and growing tips to induce a number of different physiological and morphological changes.[38]
The principal purpose of a flower is the reproduction of the individual and the species. All flowering plants are heterosporous, that is, every individual plant produces two types of spores. Microspores are produced by meiosis inside anthers and megaspores are produced inside ovules that are within an ovary. Anthers typically consist of four microsporangia and an ovule is an integumented megasporangium. Both types of spores develop into gametophytes inside sporangia. As with all heterosporous plants, the gametophytes also develop inside the spores, i. e., they are endosporic.
Flowering plants usually face evolutionary pressure to optimize the transfer of their pollen, and this is typically reflected in the morphology of the flowers and the behaviour of the plants.[49] Pollen may be transferred between plants via a number of 'vectors,' or methods. Around 80% of flowering plants make use of biotic, or living vectors. Others use abiotic, or non-living, vectors and some plants make use of multiple vectors, but most are highly specialised.[50]
Whereas in fertilization only plasmogamy, or the fusion of the whole sex cells, results, in Angiosperms (flowering plants) a process known as double fertilization, which involves both karyogamy and plasmogamy, occurs. In double fertilization the second sperm cell subsequently also enters the synergid and fuses with the two polar nuclei of the central cell. Since all three nuclei are haploid, they result in a large endosperm nucleus which is triploid.[82]
Following the pollination of a flower, fertilization, and finally the development of a seed and fruit, a mechanism is typically used to disperse the fruit away from the plant.[87] In Angiosperms (flowering plants) seeds are dispersed away from the plant so as to not force competition between the mother and the daughter plants,[88] as well as to enable the colonisation of new areas. They are often divided into two categories, though many plants fall in between or in one or more of these:[89]
Several groups of extinct gymnosperms, particularly seed ferns, have been proposed as the ancestors of flowering plants but there is no continuous fossil evidence showing exactly how flowers evolved. The apparently sudden appearance of relatively modern flowers in the fossil record posed such a problem for the theory of evolution that it was called an "abominable mystery" by Charles Darwin.
Recent DNA analysis (molecular systematics)[106] shows that Amborella trichopoda, found on the Pacific island of New Caledonia, is the only species in the sister group to the rest of the flowering plants, and morphological studies suggest that it has features which may have been characteristic of the earliest flowering plants.[107]
Besides the hard proof of flowers in or shortly before the Cretaceous,[108][109] there is some circumstantial evidence of flowers as much as 250 million years ago. A chemical used by plants to defend their flowers, oleanane, has been detected in fossil plants that old, including gigantopterids,[110] which evolved at that time and bear many of the traits of modern, flowering plants, though they are not known to be flowering plants themselves, because only their stems and prickles have been found preserved in detail; one of the earliest examples of petrification.
Many flowering plants reflect as much light as possible within the range of visible wavelengths of the pollinator the plant intends to attract. Flowers that reflect the full range of visible light are generally perceived as white by a human observer. An important feature of white flowers is that they reflect equally across the visible spectrum. While many flowering plants use white to attract pollinators, the use of color is also widespread (even within the same species). Color allows a flowering plant to be more specific about the pollinator it seeks to attract.[citation needed] The color model used by human color reproduction technology (CMYK) relies on the modulation of pigments that divide the spectrum into broad areas of absorption. Flowering plants by contrast are able to shift the transition point wavelength between absorption and reflection. If it is assumed that the visual systems of most pollinators view the visible spectrum as circular then it may be said that flowering plants produce color by absorbing the light in one region of the spectrum and reflecting the light in the other region. With CMYK, color is produced as a function of the amplitude of the broad regions of absorption. Flowering plants by contrast produce color by modifying the frequency (or rather wavelength) of the light reflected. Most flowers absorb light in the blue to yellow region of the spectrum and reflect light from the green to red region of the spectrum. For many species of flowering plant, it is the transition point that characterizes the color that they produce. Color may be modulated by shifting the transition point between absorption and reflection and in this way a flowering plant may specify which pollinator it seeks to attract.[citation needed] Some flowering plants also have a limited ability to modulate areas of absorption. This is typically not as precise as control over wavelength. Humans observers will perceive this as degrees of saturation (the amount of white in the color).
Tracheids, vessels, xylem fibres and xylem parenchyma.1. Tracheids:They are dead and without protoplasm.They are elongated or tube like cells with thick and lignified walls and tapering ends. In flowering plants, tracheids and vessels are the main water transporting elements. 041b061a72